Stilt-legged flies are a family of true flies. These slender-bodied flies are harmless, but most species resemble ichneumon or other wasps, or ants. The middle and back pairs of legs are exceptionally long and thin, while the front pair of legs are much shorter. Remember that true flies have short antennae. Members of this group, however, often walk around with the front pair of legs held forward, resembling antennae. In many species, the legs, body, or both are banded with black and yellow or orange. The forelegs, especially, may have markings that draw attention to them. The wings are narrow and in most cases are folded flat along the back. A close look at the eyes shows the characteristic large compound eyes typical of members of the true fly order (Diptera).
Some common Missouri species of stilt-legged flies include the following. None have common names.
- Calobatina geometra is mostly black but has yellowish leg bases. The two middle legs, and sometimes also the hind legs, have a black band in the middle of the yellow femur (thigh-like segment). The two forelegs have a white section near the black feet. The eyes are rusty brown and rather dark. The wings are clear.
- Rainiera antennaepes is similar to the above but has brighter red eyes, the front feet have white (not black) tips, and the black portion of the femurs is more extensive. The head is narrower. The thorax is glossy. There is a dark patch on the otherwise clear wings. The species name means "antenna-foot," referring to the forelegs that resemble antennae. Of about 20 species in its genus globally, this is the only one that occurs in North America.
- Taeniaptera trivittata has a nifty way of mimicking ants: the otherwise clear wings are marked with black dots to resemble an ant's body shape. This stilt-legged fly is mostly black except for white feet on the forelegs and some degree of rusty or yellowish coloring and sometimes subtle banding on the other legs. The eyes are dark rusty brown. The body is not glossy.
Similar species: Insects that are most likely to be confused with stilt-legged flies are usually not closely related at all: they are the insects the stilt-legged flies mimic: ichneumons and other wasps, ants, and other long-legged members of the order comprising the bees, wasps, and ants (Hymenoptera).
Habitat and Conservation
As larvae, most stilt-legged flies in our area eat decaying materials (they are saprophagous), such as rotting plant materials, decaying fungi, or manure.
As adults, stilt-legged flies in our area usually take nourishment from rotting fruits or manure.